Molecular characterization and expression analyses of toll like receptor-5 induced by Vibrio parahaemolyticus antigens in Pacific red snapper
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Toll-like receptor 5 (TLR5) is a member of TLRs family responsible for the bacterial flagellin recognition in vertebrates. Herein, the TLR5M gene structure of Pacific red snapper (Lutjanus peru) was characterized. The full-length cDNA of LpTLR5M comprises an open reading frame (ORF) of 2715 bp, encoding a polypeptide of 904 amino acids including 9 LRRs (residues 119–562) and one LRR-CT domain (residues 593–646) at the extracellular region, and a TIR domain (residues 710–904) in the cytoplasmic region. The amino acid sequence in L. peru TLR5 showed high identity (66–69%25) with TLR5 from Paralichthys olivaceus and Scophthalmus maximus. Quantitative real-time PCR (qPCR) analysis demonstrated the constitutive expression of LpTLR5M mRNA in all the examined tissues, with higher levels in intestine, liver, and head-kidney. Furthermore, expression of LpTLR5M and five cytokine genes was also investigated 24 h and one week post-stimulation in fish intraperitoneally injected with ToxA, live V. parahaemolyticus (Vp) or V. parahaemolyticus Lysate antigens. TLR5M was significantly induced in fish infected with Vp. The pro-inflammatory cytokines IL-6, IL8 and IL-12 were significantly up-regulated in head-kidney in fish stimulated with Vp, while in intestine upregulation was observed following ToxA or Lysate injection. In contrast, IL-17 mRNA was significantly up-regulated in the intestine from fish infected with live Vp at 24 h post-injection. The results indicate that Lysate and Vp antigens can induce an immune response via TLR5M and that cytokines have an important role in the defense mechanisms against V. parahaemolyticus. © 2017 Elsevier Ltd
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Inmunostimulants; Interleukins; Live Vibrio parahaemolyticus; Lysate; Pacific red snapper; TLR5M; ToxA; Vaccines amino acid; bacterial antigen; complementary DNA; elongation factor 1alpha; interleukin 10; interleukin 12; interleukin 17; interleukin 6; interleukin 8; messenger RNA; polypeptide; toll like receptor 5; bacterial antigen; fish protein; toll like receptor 5; amino acid sequence; animal experiment; animal tissue; Article; cell lysate; controlled study; cytoplasm; defense mechanism; fish; gene; gene expression; gene structure; head kidney; immune response; intestine; liver; nonhuman; nucleotide sequence; open reading frame; Pacific red snapper; Paralichthys olivaceus; protein domain; quantitative analysis; real time polymerase chain reaction; ToxA gene; turbot; upregulation; Vibrio parahaemolyticus; animal; antibody specificity; chemistry; classification; fish disease; gene expression profiling; gene expression regulation; genetics; immunology; innate immunity; Perciformes; phylogeny; physiology; randomization; sequence alignment; veterinary; Vibrio parahaemolyticus; vibriosis; Amino Acid Sequence; Animals; Antigens, Bacterial; Base Sequence; Fish Diseases; Fish Proteins; Gene Expression Profiling; Gene Expression Regulation; Immunity, Innate; Organ Specificity; Perciformes; Phylogeny; Random Allocation; Sequence Alignment; Toll-Like Receptor 5; Vibrio Infections; Vibrio parahaemolyticus
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