Food entrains clock genes but not metabolic genes in the liver of suprachiasmatic nucleus lesioned rats
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Hepatic circadian transcription, considered to be driven by the liver clock, is largely influenced by food even uncoupling it from the suprachiasmatic nucleus (SCN). In SCN lesioned rats (SCNx) we determined the influence of a physiological feeding schedule on the entrainment of clock and clock-controlled (CCG) genes in the liver. We show that clock genes and the CCG Rev-erbα and peroxisome proliferator-activated receptor alpha (PPARα) in food-scheduled intact and SCNx have a robust diurnal differential expression persisting after a 24 h fast. However, hepatic nicotinamide phosphoribosyl transferase (Nampt) shows time dependent changes that are lost in intact animals under fasting; moreover, it is unresponsive to the nutrient status in SCNx, indicating a poor reliance on liver clock genes and highlighting the relevance of SCN-derived signals for its metabolic status-related expression. © 2014 Federation of European Biochemical Societies. Published by Elsevier B.V. All rights reserved.
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Clock genes; Food-restriction; Inhibitor of DNA binding 2; Nicotinamide phosphoribosyl transferase; Peroxisome proliferator-activated receptor alpha; Sirt1 nicotinamide phosphoribosyltransferase; peroxisome proliferator activated receptor alpha; animal cell; animal experiment; animal model; animal tissue; article; body temperature; circadian rhythm; clock controlled gene; CLOCK gene; controlled study; diet restriction; food intake; gene; gene expression; ID2 gene; liver disease; liver of suprachiasmatic nucleus lesion; male; metabolic gene; metabolism; Nampt gene; neuroendocrine disease; nonhuman; nutrient; PPARalpha gene; priority journal; protein expression; rat; Rev erb alpha gene; signal transduction; suprachiasmatic nucleus; temperature fluctuation; Animalia; Rattus; Clock genes; Food-restriction; Inhibitor of DNA binding 2; Nicotinamide phosphoribosyl transferase; Peroxisome proliferator-activated receptor alpha; Sirt1; Animals; Circadian Clocks; Fasting; Food; Gene Expression Regulation; Inhibitor of Differentiation Protein 2; Liver; Male; Nicotinamide Phosphoribosyltransferase; Nuclear Receptor Subfamily 1, Group D, Member 1; PPAR alpha; Rats; Suprachiasmatic Nucleus; Temperature
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